Barrett, G. & Barrett, T. 2001

‘Cemeteries as repositories of natural and cultural diversity’, Conservation Biology, 15:6, 1820-24.


Bovyn, R., Lordon M., Grecco, A., & 2 others 2019

‘Tree cavity availability in urban cemeteries and city parks’, Journal of Urban Ecology, 5:1,1–9.


Buchholz, S., Blick, T., Hannig, K. & 8 others 2016

‘Biological richness of a large urban cemetery in Berlin. Results of a multi-taxon approach’, Biodiversity Data Journal, 4, e7057.

Čanády, A. & Mošanský, L. 2017

‘Public Cemetery as a biodiversity hotspot for birds and mammals in the urban environment of Kosice City (Slovakia)’, Zoology and Ecology, 27: 3-4, 185-95.


Czarna, A. 2016

‘Vascular plant flora in the Cytadela cemeteries in Poznań (Poland)’, Acta Agrobotanica 69:4, 1695.


de Lacy, P. & Shackleton, C. 2017

Woody plant species richness, composition and structure in urban sacred sites, Grahamstown, South Africa’, Urban Ecosystems, 20, 1169-1179.

Długozima, A. & Kosiacka-Beck, E. 2020

‘How to enhance the environmental impact of contemporary cemeteries an in urban context’, Sustainability, 12:6, 2374.

Długozima, A. & Nejman, R. 2022

‘Attractiveness of cemeteries versus socioeconomic and spatial development of non-metropolitan cities in Poland’, Acta Scientiarum Polonorum Administratio Locorum 21:1, 83-104.

Gilbert, O. 1989

‘Cemeteries’, in O. Gilbert (ed.) The Ecology of Urban Habitats, Springer Netherlands, 218-38.

Holden, M. & McDonald-Madden, E. 2017

‘Conservation from the grave: human burials to fund the conservation of threatened species’, Conservation Letters, 11:1, e12421.

Jaganmohan, M., Vailshery L., Mundoli S. & Nagendra H. 2018

‘Biodiversity in sacred urban spaces of Bengaluru, India’, Urban Forestry and Urban Greening, 32, 64–70.


Kamran, S., Khan, S., Ahrmad, Z. & 3 others . 2020

‘The role of graveyards in species conservation and beta diversity: a vegetation appraisal of sacred habitats from Bannu, Pakistan’, Journal of Forestry Research, 31, 1147-1158.


Konic, J., Essl, F. & Lenzner, B. 2021

‘To care or not to care? Which factors influence the distribution of early-flowering geophytes at the Vienna Central Cemetery’, Sustainability, 13, 4657.


Kowirak, I., Buchholz, S., von der Lippe, M. & Seitz, B. 2016

‘Biodiversity functions of urban cemeteries: evidence from one of the largest Jewish cemeteries in Europe’, Urban Forestry & Urban Greening, 19, 68-78.


Laske, D. 1994

‘Cemeteries: ecological niches in populated areas’, Naturwissenschaften, 81, 218-223.


Löki, V., Deák, B., Lukács, A.B., Molnár, A. 2019

‘Biodiversity potential of burial places: a review on the flora and fauna of cemeteries and churchyards’, Global Ecology and Conservation, 18, 1-14.


Löki, V., Molnár, A. Süveetes, K. & 8 others 2019

‘Predictors of conservation value of Turkish cemeteries: a case study using orchids’, Landscape and Urban Planning, 186, 36-44.


Löki, V., Schmotzer, A., Takács, A. & 5 others 2020

‘The protected flora of long‐established cemeteries in Hungary: using historical maps in biodiversity conservation’, Ecology and Evolution, 10:14, 7497-7508.


McPherson, E.& Nilon, C. 1987

‘A habitat suitability index model for gray squirrel in an urban cemetery’, Landscape Journal, 6, 21–30.


Molnár, A., Takács, A., Mizsei, E. & 3 others 2017

‘Religous difference affecting orchid diversity of Albanian graveyards’, Pakistani Journal of Botany, 49:1, 289-303.



The Cemetery Research Group runs two events a year: in May and in November. Follow the links and send in an abstract